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However, this hypothesis has been challenged bysubsequent research. At present, it is commonly believed that liverstem/progenitor cells are the potential source of HCC, intrahepaticcholangiocarcinoma (ICC), combined hepatocellularcholangiocarcinoma (CHC) and cholangiolocellular carcinoma (CLC), asubtype of cholangiocellular carcinoma (CC) (–).To study the effect of oval cells upon tumorigenesis, de Lima () established a rat modelof non-alcoholic steatohepatitis (NASH), cirrhosis and HCC, showingthat oval cells could proliferate in this model and that thesecells may be the origin of malignancy. In another model (theSolt-Farber carcinogenic model), hepatic progenitor cells,identified by the expression of glypican-3 (GPC3), were shown toplay an important role in hepatic carcinogenesis (). A summary of the origin of LCSCs ispresented in .